Wright-Fisher model with negative mutation rates ABSTRACT: An n-leaf cladogram is an unrooted tree with n leaves and every internal vertex of degree three. David Aldous introduced a natural Markov chain on the space of Cladograms that consists of picking a leaf at random and randomly re-attaching it to one of the remaining edges. As n grows to infinity this diffusion, suitably rescaled in time, is conjectured to converge to a diffusion on the space of continuum trees. We talk about the local structure of the limiting diffusion seen from the point of view of finitely many branch points. Locally it is a finite dimensional diffusion with state space in the unit simplex and is not reversible. However we show how it is related to another famous Markov chain model, the Wright-Fisher model, which consists of sampling balls of different colors from one urn to fill another. The key seems to be interpreting these diffusions as functions of Galton-Watson trees with immigration. As a benefit of working with diffusions explicit calculations are easy to derive. BIO: Soumik Pal did his undergraduate at the Indian Statistical Institute in Kolkata, and got his Ph.D. at the statistics department of Columbia University, under the supervision of I. Karatzas in 2006. After a postdoc at Cornell, he became a professor in the math department at UW in 2008.